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6H 2 O + 6CO 2 ----------> C 6 H 12 O 6 + 6O 2

carotenechlorophyll

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Слайд 16H2O + 6CO2 ----------> C6H12O6+ 6O2

6H2O + 6CO2 ----------> C6H12O6+ 6O2

Слайд 3carotene
chlorophyll

carotenechlorophyll

Слайд 10Electron micrograph of 2D crystals of the LH1-Reaction centre photosynthetic

unit.

Electron micrograph of 2D crystals of the LH1-Reaction centre photosynthetic unit.

Слайд 18Thylakoid membrane

Thylakoid membrane

Слайд 19The "Z scheme"

The

Слайд 24Схема световой стадии фотосинтеза высших растений:
I - комплекс фотосистемы ФС

I;
II - комплекс ФС II;
III - цитохром-ный b6-

f-комплекс;
CF1-CF0- сопрягающий комплекс;
Фд - ферредоксин, ФП - флавопротеиновая Фд - НАДФ-редуктаза;
Qa - Qb - вторичные акцепторы хинонной природы.
Схема световой стадии фотосинтеза высших растений:I - комплекс фотосистемы ФС I; II - комплекс ФС II; III

Слайд 26To avoid an over-acidification of the thylakoid lumen and the

detrimental production of reactive oxygen species, the photosynthetic ATP and

NADPH production is closely adjusted to the ATP and NADPH consumption rates by the Calvin cycle and the subsequent reactions of dark metabolism.

High rates of ATP and NADPH production in young leaves with high assimilation capacity are facilitated by high contents of the rate-limiting components cytochrome b6f complex, plastocyanin and ATP synthase. In response to a diminished assimilation capacity, ageing leaves as well as mutants with an impaired Calvin cycle repress photosynthetic electron transport via down-regulation of the cytochrome b6f complex, plastocyanin and ATP synthase. The contents of both photosystems remain largely unaltered, independent of the metabolic demands of the leaf.

To avoid an over-acidification of the thylakoid lumen and the detrimental production of reactive oxygen species, the

Слайд 27Reversible phosphorylation of LHCII. When PSII is overexcited relative to

PSI, plastoquinol (PQH2) accumulates. A high level of plastoquinol activates

a protein kinase that phosphorylates LHCII. Addition of a phosphate group weakens the interaction between LHCII and the PSII core antenna, causing LHCII to dissociate from PSII. The input of light to PSII is diminished and PSII slows down, thus allowing PSI to oxidize excess PQH2 to plastoquinone (PQ), which, in turn, deactivates the protein kinase. LHCII is dephosphorylated by a protein phosphatase, allowing LCHII to reform in association with PSII.
Reversible phosphorylation of LHCII.   When PSII is overexcited relative to PSI, plastoquinol (PQH2) accumulates. A

Слайд 30The first steps in the Calvin ccycle

The first steps in the Calvin ccycle

Слайд 31Calvin cycle

Calvin cycle

Слайд 34Hatch-Slack pathway

Hatch-Slack pathway

Слайд 35The three variants of the C4 photosynthetic carbon cycle.
The

variants differ principally in
(1) the nature of the four-carbon

acid (malate or aspartate) transported into the bundle sheath cell and of the three-carbon acid (pyruvate or alanine) returned to the mesophyll cell and
(2) the nature of the enzyme that catalyzes the decarboxylation step in the bundle sheath cell. The three variants are named after the enzymes that catalyze the decarboxylation reactions. Representatives of each variant include
- maize, crabgrass, sugarcane, sorghum (NADP malic enzyme);
- pigweed, millet (NAD malic enzyme);
- guinea grass (phosphoenolpyruvate carboxykinase).
The three variants of the C4 photosynthetic carbon cycle. The variants differ principally in (1) the nature

Слайд 36Supplementary energy required for the operation of C4 photosynthesis

Supplementary energy required for the operation of C4 photosynthesis

Слайд 37Leaf anatomy of a C3 (top) and C4 (bottom) plant

Leaf anatomy of a C3 (top) and C4 (bottom) plant

Слайд 39Supplementary energy required for the operation of CAM plants

Supplementary energy required for the operation of CAM plants

Слайд 40RuBisCo

RuBisCo

Слайд 42The cyanobacterial CO2 concentrating mechanism.

The cyanobacterial CO2 concentrating mechanism.

Слайд 44Photorespiration

Photorespiration

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